Can't get LDAP to work

I have created an ldap-config.properties file as follows:

java.naming.provider.url=ldaps://ldap.jumpcloud.com:636/O=<SECRET>,DC=jumpcloud,DC=com
java.naming.security.principal=bind_dn
java.naming.security.credentials=<SECRET>
teamcity.users.base=CN=users
teamcity.users.login.filter=(&(objectClass=person)(memberOf=cn=Teamcity,ou=Users,o=<SECRET>,dc=jumpcloud,dc=com))
teamcity.users.username=uid
teamcity.users.login.filter=(&(objectClass=person)(memberOf=cn=Teamcity,ou=Users,o=<SECRET>,dc=jumpcloud,dc=com))
teamcity.options.users.synchronize=true
teamcity.options.groups.synchronize=false
teamcity.options.createUsers=true
teamcity.options.deleteUsers=true
teamcity.options.syncTimeout=3600000
teamcity.groups.property.member=member
teamcity.users.property.displayName=displayName
teamcity.users.property.email=mail

I when I switched the auth mechanism to LDAP and tried to log in with my LDAP user account, I got the following error:

[2018-05-18 19:37:22,651] INFO - jetbrains.buildServer.LDAP -
Starting synchronization session
[2018-05-18 19:37:22,651] INFO - jetbrains.buildServer.LDAP -
Fetching remote users and groups
[2018-05-18 19:37:23,125] WARN - jetbrains.buildServer.LDAP - 
Error while retrieving LDAP users, skipping users 
synchronization: LDAP search operation returned an error 
while retrieving users. While initializing LDAP connection. 
LDAP server says: Invalid DN syntax. 
Original error: org.springframework.ldap.InvalidNameException:
[LDAP: error code 34 - invalid DN]; nested exception is 
javax.naming.InvalidNameException: [LDAP: error code 34 - invalid DN]
[2018-05-18 19:37:23,125] INFO - jetbrains.buildServer.LDAP - Skipping 
groups synchronization as 'teamcity.options.groups.synchronize' 
property is not set to 'true'
[2018-05-18 19:37:23,125] INFO - jetbrains.buildServer.LDAP - Last 
synchronization statistics: user sync enabled=false, created users=0, 
updated users=0, deleted users=0, remote users=0, matched users=0, group 
sync enabled=false, created groups=0, updated groups=0, deleted groups=0, 
remote groups=0, matched groups=0, duration=474ms, errors=1, errors: 
[Error while retrieving LDAP users, skipping users synchronization: 
LDAP search operation returned an error while retrieving users. While 
initializing LDAP connection. LDAP server says: Invalid DN syntax. 
Original error: org.springframework.ldap.InvalidNameException: [LDAP: 
error code 34 - invalid DN]; nested exception is 
javax.naming.InvalidNameException: [LDAP: error code 34 - invalid DN]]

I don't even see where a DN is specified in the config above.  How do I resolve this error?  Does anyone have a non-AD LDAP configuration that is working?  I was able to do an ldapsearch successfully with this configuration:

  ldapsearch -H ldaps://ldap.jumpcloud.com:636 \
  -b u=Users,o=<SECRET>,dc=jumpcloud,dc=com \
  -D uid=bind_dn,ou=Users,o=<SECRET>,dc=jumpcloud,dc=com \
  -w <SECRET> \
  (&(objectClass=person)(memberOf=cn=Teamcity,ou=Users,o=<SECRET>,dc=jumpcloud,dc=com))

This returns a list of members of the Teamcity group exactly as expected